(born 1966) is a professor of anatomy at the University of Tampere, Finland.
He graduated from the University of Oulu in 1991 (M.D.) and obtained his PhD-degree from the same university in 1994.
In 1996-1998 he worked as a visiting researcher at Saint Louis University.
In 2002 he moved to the University of Tampere where he started to work as a professor of medical technology and biotechnology and later in 2008 he was appointed to the post of professor in anatomy.
His research work has been focused on pH-regulation, carbonic anhydrases and regulation of iron homeostasis.